Hae opetus/tentit
ASPB1102 Arctic Ecosystems and Adaptation of Species to Arctic Environment, 5 op 
Koodi ASPB1102  Voimassaolo 01.01.1950 -
Nimi Arctic Ecosystems and Adaptation of Species to Arctic Environment  Lyhenne Arctic Ecosyste 
Laajuus5 op  Vanhenemisaika  
TyyppiPerusopinnot Oppiaine0959 Monialainen, kansainvälinen 
LajiOpintojakso  Tuntimäärä  
Opinto-oikeus   ArvosteluHyväksytty, hylätty, arvosanat 1-5 
Suositeltu suoritusaika 
Vastuuyksikkö Yhteiskuntatieteiden tiedekunta 

Mari Kuoppamaa 
Osmo Rätti 
Päivi Soppela 
Sari Stark 
Minna Turunen 


After completion of the course the student is able to

  • identify the main features and functions of circumpolar tundra, boreal forest and marine ecosystems
  • recognize interactions and relationships between the Arctic ecosystems and other parts of the globe
  • examine evolution and adaptation mechanisms of plants and animals into Arctic environment.

The course contains the following themes:

1) The evolution and distribution of tundra and boreal forest ecosystems, emphasizing vegetation, plant-animal interactions and natural and anthropogenic disturbance patterns and processes.

2) Processes and global interactions affecting to marine productivity and biodiversity, interactions between marine and terrestrial ecosystems and threats to Arctic marine life.

3) Morphological, physiological and behavioural adaptations of plants and/or animals to the northern environment with basic information on evolution, natural selection and challenges of the climate change.

Edeltävät opinnot 

Course ASPB1101 Introduction to the Arctic (5 ECTS) has to be passed before attending this course.

Toteutus ja työmuodot 

Lectures (30 h), independent work (100 h).

Vaadittavat suoritukset 

Active participation in the lectures, successful completion of a written essay or an exam.


Literature (suggested):

ACIA 2005. Arctic Climate Impact Assessment. Scientific report. http://www.acia.uaf.edu/pages/scientific.html. Chapters 7 & 14.

Arctic Marine Biodiversity Monitoring Plan, pp. 108–125. Available at www.caff.is

Barboza, P. S., Parker, K. L. & Hume, I.D. 2010. Integrative Wildlife Nutrition. Springer Verlag, 342 pp.

Bardgett, R. 2005. The Biology of Soil. A community and ecosystem approach. Oxford University Press, Chapter 5. Above-ground trophic interactions and soil biological communities (pp. 119–139).

Bigras F. J., Colombo S. J. 2001. Conifer Cold Hardiness. Kluwer Academic Publishers. Dordrecht Boston London. 596 pp.

Blix, A. S. 2005. Arctic Animals and Their Adaptations to Life on the Edge. Tapir Academic Press.

Blix, A. S. 2016. Adaptations to polar life in mammals and birds. J. Exp. Biol. 219:1093-1105. doi: 10.1242/jeb.120477.

CAFF (Conservation of Arctic Flora and Fauna) 2001: Arctic Flora and Fauna: Status and Conservation. Edita. 272 pp.

Cockburn, A. 1991. An Introduction to Evolutionary Ecology. Blackwell, Oxford.

Crawford R. M. M. 1989. Studies in Plant Survival. Blackwell Scientific Publications. Oxford. 296p.

Crawford R. M. M. 2008. Plants at the Margin. Ecological Limits and Climate Change. Cambridge University Press.

Crawford, R. M. M. 2014. Tundra-taiga biology. Human, Plant and Animal Survival in the Arctic. Oxford University Press, 270 pp.

Davenport, J. 1992. Animal life at low temperature. Chapman & Hall, London.

Eamer, J., Donaldson, G. M., Gaston, H. et al. 2013. Life linked to Ice. A guide to sea-ice associated biodiversity in this time of rapid change. CAFF Assessment Series No. 10. Coservation of Arctic Flora and Fauna, Iceland. Available at: target=_blank>http://www.caff.is/sea-ice-associated-biodiversity/sea-ice-publications

Glossary of terms for thermal physiology, 2001. Third edition. The Japanese Journal of Physiology. 51: 2. Available at target=_blank>http://www.or.org/pdf/ThermalPhysiologyGlossary.pdf

IPCC 2014. Chapter 28. Polar Regions. Observed changes, Terrestrial ecosystems, Arctic (pp. 1577–1581), Projected Impacts, Terrestrial ecosystems, Arctic (pp. 1589–1590).

Marchand P. J. 1991. Life in the Cold. An Introduction to Winter Ecology. University Press of New England. Hanover and London. 239 pp.

Marchand, P. J. 1996. Life in the cold: An introduction to winter ecology. University Press of New England, Hanover.

Pond, C. 1998. Fats of life. Cambridge University Press. 337 pp.

Sakai A., Larcher W. 1987. Frost Survival of Plants. Responses and Adaptation to freezing stress. Ecological Studies 62. Springer-Verlag. 321 pp.

Skelton, P. (ed.) 1992. Evolution: a biological and palaeontological approach (Book 1). The Open University.


Grades 1-5/fail


Autumn / 1 st period


4 Oct time 12-16 Lecture Hall LS1 (= Fellman-sali), University Main Building, Yliopistonkatu 8


Principle tutor:
University Researcher Sari Stark, PhD, Arctic Centre.

University Researcher Osmo Rätti, PhD; University Researcher Päivi Soppela, PhD; University Researcher Minna Turunen, PhD; University Researcher Sari Stark, PhD; Research Professor Bruce Forbes, PhD (Arctic Centre).




This course is compulsory for the students of the Arctic Studies Programme and optional for other students.


Meneillä oleva ja tuleva opetus
Toiminnot Nimi Tyyppi op Opettaja Aikataulu
ilmoittautumisaika on päättynyt Arctic Ecosystems and Adaptation of Species to Arctic Environment  Luentokurssi  Mari Kuoppamaa,
Osmo Rätti,
Päivi Soppela,
Sari Stark,
Minna Turunen 
23.09.19ma 12.00-15.00
24.09.19ti 12.00-15.00
25.09.19ke 09.00-12.00
26.09.19to 09.00-12.00
27.09.19pe 09.00-12.00
30.09.19ma 09.00-12.00
01.10.19ti 12.00-15.00
02.10.19ke 12.00-15.00
03.10.19to 12.00-15.00
07.10.19ma 12.00-15.00

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